Wesley, 34, and Tonkin, 27, who were last seen in public together on Dec. 20, first met on the set of The Vampire Diaries in 2012, and started dating the following summer. Tonkin had a recurring role on the fourth season of the CW hit, before her character joined The Originals spin off in 2013.
"They are still good friends. The relationship just ran its course," a source tells E! News, who was first to break the news.