Dakota Johnson and Matthew Hitt Split After Two Years of Dating

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The summer of splits continues.

The 26-year-old actress split with her on-again, off-again boyfriend of two years, 29-year-old British musician Matthew Hitt, a source confirms to ET. The two first stepped out as a couple in July 2014, but broke up in February 2015. A few months later, they had rekindled the relationship, showing PDA in New York City.

Ironically, the Fifty Shades of Grey star's last film was the comedy, How to Be Single. When ET caught up Johnson in February, she shared the sound dating advice she got from her famous mother, actress Melanie Griffith, when she was younger.

WATCH: Dakota Johnson and Jamie Dornan Get Hot and Heavy Filming Iconic 'Fifty Shades Darker' Scene

"She always said not to compromise myself or my work or my dreams for anyone," Johnson recalled. "Especially a dude."

Johnson is currently filming the Fifty Shades trilogy in Vancouver, Canada. In April, her and co-star Jamie Dornan -- who plays billionaire Christian Grey -- sported wedding rings on set.

... And there's also been plenty of smooching.

WATCH: Dakota Johnson Accidentally Drops the F-Bomb on 'The Tonight Show'

Though, according to Johnson herself, she's totally "over" filming her more racier Fifty Shades scenes.

"Well, we're not having actual sex," she told Interview magazine in April. "But I've been simulating sex for seven hours straight right now, and I'm over it."

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